Quarterly
Annual
| Unit: USD | 2024-03-31 | 2023-12-31 | 2023-09-30 | 2023-06-30 | 2023-03-31 | 2022-12-31 | 2022-09-30 | 2022-06-30 | 2022-03-31 | 2021-12-31 | 2021-09-30 | 2021-06-30 | 2021-03-31 | 2020-12-31 | 2020-09-30 | 2020-06-30 | 2020-03-31 | 2019-12-31 | 2019-09-30 | 2019-06-30 | 2019-03-31 | 2018-12-31 | 2018-09-30 | 2018-06-30 | 2018-03-31 | 2017-12-31 | 2017-09-30 | 2017-06-30 | 2017-03-31 | 2016-12-31 | 2016-09-30 | 2016-06-30 | 2016-03-31 | 2015-12-31 | 2015-09-30 | 2015-06-30 |
|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
assets | ||||||||||||||||||||||||||||||||||||
current assets: | ||||||||||||||||||||||||||||||||||||
cash and cash equivalents | 33,015,000 | 43,921,000 | 18,981,000 | 34,076,000 | 14,465,000 | 13,376,000 | 126,625,000 | 24,360,000 | 34,889,000 | 67,907,000 | 76,099,000 | 18,813,000 | 12,629,000 | 34,959,000 | 141,075,000 | 88,220,000 | 24,446,000 | 16,123,000 | 17,395,000 | 15,468,000 | 41,437,000 | 10,711,000 | 11,473,000 | 3,569,000 | 7,446,000 | 8,000,000 | 11,982,000 | 27,548,000 | 12,929,000 | 16,168,000 | 15,416,000 | 16,030,000 | 24,991,000 | 57,555,000 | 100,521,000 | |
short-term investments | 272,646,000 | 283,491,000 | 174,472,000 | 169,724,000 | 180,460,000 | 224,265,000 | 122,237,000 | 143,500,000 | 136,980,000 | 94,396,000 | 104,529,000 | 72,247,000 | 84,185,000 | 70,622,000 | 1,998,000 | 11,237,000 | 24,397,000 | 29,193,000 | 40,097,000 | 22,519,000 | 41,592,000 | 50,509,000 | 66,376,000 | 69,206,000 | 73,240,000 | 75,219,000 | ||||||||||
accounts receivable | 42,000 | 167,000 | 219,000 | 284,000 | 668,000 | 392,000 | 382,000 | 1,317,000 | 132,000 | 25,000,000 | 45,343,000 | |||||||||||||||||||||||||
prepaid expenses and other current assets | 2,548,000 | 2,455,000 | 3,743,000 | 3,237,000 | 3,191,000 | 2,960,000 | 3,995,000 | 3,112,000 | 4,095,000 | 4,710,000 | 3,121,000 | 3,333,000 | 3,668,000 | 1,520,000 | 1,475,000 | 1,018,000 | 2,273,000 | 1,650,000 | 5,734,000 | 1,657,000 | 939,000 | 1,242,000 | 1,098,000 | 1,850,000 | 1,335,000 | 1,308,000 | 884,000 | 129,000 | 263,000 | 843,000 | 480,000 | 501,000 | 432,000 | 996,000 | 569,000 | |
total current assets | 308,251,000 | 330,034,000 | 197,415,000 | 207,321,000 | 198,784,000 | 240,993,000 | 253,239,000 | 172,289,000 | 176,096,000 | 192,013,000 | 183,749,000 | 139,736,000 | 100,482,000 | 107,101,000 | 142,550,000 | 91,236,000 | 38,088,000 | 42,302,000 | 52,454,000 | 57,222,000 | 64,895,000 | 53,545,000 | 63,080,000 | 71,795,000 | 77,987,000 | 82,548,000 | 88,085,000 | 49,626,000 | 52,937,000 | 67,030,000 | 74,214,000 | 80,713,000 | 87,686,000 | 93,647,000 | 101,090,000 | |
restricted cash, noncurrent | 269,000 | 268,000 | 254,000 | 254,000 | 254,000 | 254,000 | 254,000 | 254,000 | 254,000 | 254,000 | 254,000 | 254,000 | 254,000 | 254,000 | 254,000 | 254,000 | 254,000 | 254,000 | 254,000 | |||||||||||||||||
property and equipment | 1,356,000 | 1,484,000 | 1,533,000 | 1,672,000 | 1,689,000 | 1,584,000 | 1,608,000 | 1,596,000 | 1,728,000 | 1,716,000 | 1,372,000 | 1,527,000 | 1,674,000 | 1,785,000 | 1,695,000 | 1,531,000 | 1,669,000 | 1,552,000 | 1,201,000 | 1,176,000 | 1,221,000 | 1,196,000 | 1,296,000 | 1,197,000 | 1,127,000 | 1,089,000 | 1,116,000 | |||||||||
operating lease, right-of-use asset | 7,317,000 | 7,510,000 | 7,697,000 | 7,877,000 | 8,051,000 | 8,219,000 | 8,381,000 | 8,539,000 | 8,691,000 | 8,837,000 | 8,980,000 | 9,118,000 | 9,251,000 | 9,401,000 | 9,534,000 | 9,674,000 | 9,838,000 | 9,985,000 | 10,830,000 | 11,303,000 | 603,000 | |||||||||||||||
long-term investments | 56,453,000 | 40,556,000 | 33,502,000 | 35,561,000 | 53,170,000 | 35,481,000 | 28,016,000 | 33,041,000 | 46,948,000 | 52,866,000 | 39,064,000 | 9,120,000 | 18,316,000 | 25,549,000 | ||||||||||||||||||||||
total assets | 373,646,000 | 379,852,000 | 240,547,000 | 252,836,000 | 262,100,000 | 286,686,000 | 291,599,000 | 215,826,000 | 233,938,000 | 255,900,000 | 233,480,000 | 159,818,000 | 129,977,000 | 144,090,000 | 154,033,000 | 102,695,000 | 49,849,000 | 54,093,000 | 64,739,000 | 70,087,000 | 67,105,000 | 54,873,000 | 64,508,000 | 73,124,000 | 80,699,000 | 85,222,000 | 90,786,000 | 49,644,000 | 53,009,000 | 61,935,000 | 67,342,000 | 74,565,000 | 80,937,000 | 87,909,000 | 93,842,000 | 101,247,000 |
liabilities and stockholders’ equity | ||||||||||||||||||||||||||||||||||||
current liabilities: | ||||||||||||||||||||||||||||||||||||
accounts payable | 4,170,000 | 3,593,000 | 3,068,000 | 1,645,000 | 1,662,000 | 4,286,000 | 3,240,000 | 1,812,000 | 1,903,000 | 3,349,000 | 1,839,000 | 1,393,000 | 466,000 | 582,000 | 742,000 | 548,000 | 468,000 | 1,543,000 | 5,778,000 | 2,399,000 | 743,000 | 1,716,000 | 440,000 | 750,000 | 528,000 | 193,000 | 1,076,000 | 815,000 | 1,278,000 | 1,012,000 | 2,110,000 | 1,196,000 | 994,000 | 1,552,000 | 3,432,000 | |
accrued liabilities | 16,891,000 | 21,187,000 | 17,788,000 | 13,684,000 | 12,030,000 | 14,003,000 | 13,744,000 | 12,525,000 | 9,203,000 | 9,417,000 | 8,653,000 | 4,614,000 | 3,953,000 | 5,777,000 | 5,065,000 | 3,824,000 | 2,551,000 | 5,285,000 | 5,825,000 | 5,518,000 | 5,371,000 | 4,277,000 | 4,314,000 | 585,000 | 586,000 | 382,000 | 642,000 | |||||||||
deferred revenue, current | 10,227,000 | 16,288,000 | 30,092,000 | 37,252,000 | 40,401,000 | 35,571,000 | 46,210,000 | 42,917,000 | 51,760,000 | 51,773,000 | 48,100,000 | 48,957,000 | 34,522,000 | 31,627,000 | 28,505,000 | 23,688,000 | ||||||||||||||||||||
operating lease liability, current | 950,000 | 912,000 | 870,000 | 829,000 | 790,000 | 756,000 | 718,000 | 682,000 | 647,000 | 617,000 | 747,000 | 715,000 | 684,000 | 655,000 | 219,000 | 452,000 | ||||||||||||||||||||
total current liabilities | 32,238,000 | 41,980,000 | 51,818,000 | 53,410,000 | 57,089,000 | 57,996,000 | 68,459,000 | 62,618,000 | 68,165,000 | 69,778,000 | 63,933,000 | 60,246,000 | 43,367,000 | 41,167,000 | 35,849,000 | 28,179,000 | 5,105,000 | 8,681,000 | 13,672,000 | 12,556,000 | 8,848,000 | 8,127,000 | 6,867,000 | 3,426,000 | 2,928,000 | 1,895,000 | 2,724,000 | 3,578,000 | 3,789,000 | 5,238,000 | 5,873,000 | 4,578,000 | 4,419,000 | 4,465,000 | 6,123,000 | |
deferred revenue, noncurrent | 929,000 | 17,812,000 | 20,475,000 | 25,636,000 | 39,185,000 | 30,803,000 | 42,083,000 | 37,346,000 | 50,830,000 | 30,961,000 | 38,621,000 | 15,250,000 | 21,348,000 | 29,988,000 | 36,468,000 | |||||||||||||||||||||
operating lease liability, noncurrent | 8,705,000 | 9,002,000 | 9,286,000 | 9,563,000 | 9,832,000 | 10,085,000 | 10,328,000 | 10,564,000 | 10,794,000 | 11,009,000 | 11,214,000 | 11,413,000 | 11,608,000 | 11,815,000 | 11,987,000 | 11,947,000 | 11,716,000 | 11,429,000 | 11,350,000 | 10,919,000 | ||||||||||||||||
total liabilities | 40,943,000 | 51,911,000 | 78,916,000 | 83,448,000 | 92,557,000 | 107,266,000 | 109,590,000 | 116,290,000 | 118,511,000 | 134,997,000 | 110,655,000 | 115,987,000 | 76,683,000 | 81,932,000 | 86,565,000 | 86,467,000 | 24,990,000 | 24,619,000 | 29,908,000 | 24,666,000 | 10,522,000 | 10,282,000 | 9,524,000 | 6,577,000 | 6,818,000 | 6,305,000 | 7,814,000 | 6,499,000 | ||||||||
commitments and contingencies | ||||||||||||||||||||||||||||||||||||
stockholders’ equity: | ||||||||||||||||||||||||||||||||||||
preferred stock, 0.001 par value per share; 10,000,000 shares authorized at march 31, 2024 and december 31, 2023; zero shares issued and outstanding at march 31, 2024 and december 31, 2023 | ||||||||||||||||||||||||||||||||||||
common stock | 66,000 | 60,000 | 49,000 | 49,000 | 48,000 | 46,000 | 44,000 | 30,000 | 30,000 | 30,000 | 29,000 | 24,000 | 24,000 | 24,000 | 24,000 | 19,000 | 19,000 | 19,000 | 19,000 | 19,000 | 19,000 | 14,000 | 14,000 | 14,000 | 14,000 | 14,000 | 1,000 | 16,000 | 16,000 | 16,000 | 15,000 | 15,000 | 15,000 | 15,000 | 15,000 | |
additional paid-in capital | 606,963,000 | 583,629,000 | 424,227,000 | 420,372,000 | 407,129,000 | 404,456,000 | 388,731,000 | 292,699,000 | 290,184,000 | 287,345,000 | 273,813,000 | 181,300,000 | 179,746,000 | 177,947,000 | 177,009,000 | 119,651,000 | 118,417,000 | 117,371,000 | 116,591,000 | 115,704,000 | 115,031,000 | 90,664,000 | 89,933,000 | 89,387,000 | 88,864,000 | 88,346,000 | 88,049,000 | 238,980,000 | 238,434,000 | 234,670,000 | 233,808,000 | 233,016,000 | 232,309,000 | 231,621,000 | 231,294,000 | |
accumulated other comprehensive income | -263,000 | -14,000 | 5,000 | 23,000 | 3,000 | 3,000 | 15,000 | |||||||||||||||||||||||||||||
accumulated deficit | -274,063,000 | -256,145,000 | -262,105,000 | -250,383,000 | -237,227,000 | -223,961,000 | -205,097,000 | -191,824,000 | -173,726,000 | -166,199,000 | -151,032,000 | -137,538,000 | -126,509,000 | -115,866,000 | -109,524,000 | -103,387,000 | -93,459,000 | -87,926,000 | -81,786,000 | -70,311,000 | -58,455,000 | -46,074,000 | -34,938,000 | -22,799,000 | -14,892,000 | -9,384,000 | -5,066,000 | -192,926,000 | -189,216,000 | -180,300,000 | -172,587,000 | -165,154,000 | -156,675,000 | -148,837,000 | -142,274,000 | -136,185,000 |
total stockholders’ equity | 332,703,000 | 327,941,000 | 161,631,000 | 169,388,000 | 169,543,000 | 179,420,000 | 182,009,000 | 99,536,000 | 115,427,000 | 120,903,000 | 122,825,000 | 43,831,000 | 53,294,000 | 62,158,000 | 67,468,000 | 16,228,000 | 24,859,000 | 29,474,000 | 34,831,000 | 45,421,000 | 56,583,000 | 44,591,000 | 54,984,000 | 66,547,000 | 73,881,000 | 82,972,000 | 46,066,000 | 49,220,000 | 62,104,000 | 68,692,000 | 76,359,000 | 83,490,000 | 89,377,000 | 95,124,000 | ||
total liabilities and stockholders’ equity | 373,646,000 | 379,852,000 | 240,547,000 | 252,836,000 | 262,100,000 | 286,686,000 | 291,599,000 | 215,826,000 | 233,938,000 | 255,900,000 | 233,480,000 | 159,818,000 | 129,977,000 | 144,090,000 | 154,033,000 | 102,695,000 | 49,849,000 | 54,093,000 | 64,739,000 | 70,087,000 | 67,105,000 | 49,644,000 | 53,009,000 | 67,342,000 | 74,565,000 | 80,937,000 | 87,909,000 | 93,842,000 | 101,247,000 | |||||||
deferred tax asset | 146,000 | 151,000 | 152,000 | 155,000 | 101,000 | 107,000 | 221,000 | 214,000 | 61,000 | 63,000 | ||||||||||||||||||||||||||
current portion of long-term debt | 2,206,000 | 3,380,000 | 4,547,000 | 4,682,000 | 4,652,000 | 4,622,000 | 4,594,000 | 4,567,000 | 3,742,000 | 2,526,000 | 1,318,000 | 119,000 | 1,742,000 | 418,000 | 2,079,000 | 2,064,000 | 2,048,000 | 2,034,000 | 2,018,000 | 1,508,000 | 995,000 | 490,000 | ||||||||||||||
preferred stock, 0.001 par value per share; 10,000,000 shares authorized at december 31, 2023 and 2022; zero shares issued and outstanding at december 31, 2023 and 2022 | ||||||||||||||||||||||||||||||||||||
treasury stock | ||||||||||||||||||||||||||||||||||||
accumulated other comprehensive gain | 397,000 | -273,000 | 15,000 | 45,000 | 33,000 | 53,000 | -41,000 | -55,000 | -118,000 | 10,000 | 7,000 | 9,000 | ||||||||||||||||||||||||
preferred stock, 0.001 par value per share; 10,000,000 shares authorized at september 30, 2023 and december 31, 2022; zero shares issued and outstanding at september 30, 2023 and december 31, 2022 | ||||||||||||||||||||||||||||||||||||
accumulated other comprehensive loss | -540,000 | -650,000 | -407,000 | -1,121,000 | -1,669,000 | -1,369,000 | -1,061,000 | -12,000 | -13,000 | -25,000 | -55,000 | -105,000 | -59,000 | -12,000 | ||||||||||||||||||||||
preferred stock, 0.001 par value per share; 10,000,000 shares authorized at june 30, 2023 and december 31, 2022; zero shares issued and outstanding at june 30, 2023 and december 31, 2022 | ||||||||||||||||||||||||||||||||||||
preferred stock, 0.001 par value per share; 10,000,000 shares authorized at march 31, 2023 and december 31, 2022; zero shares issued and outstanding at march 31, 2023 and december 31, 2022 | ||||||||||||||||||||||||||||||||||||
long-term debt | 1,025,000 | 2,206,000 | 3,380,000 | 4,547,000 | 5,707,000 | 6,458,000 | 7,602,000 | 8,741,000 | 9,873,000 | 8,169,000 | 4,509,000 | 4,886,000 | 1,191,000 | 1,674,000 | 2,155,000 | 2,631,000 | 3,104,000 | 3,570,000 | 4,039,000 | 4,500,000 | ||||||||||||||||
preferred stock, 0.001 par value per share; 10,000,000 shares authorized at december 31, 2022 and 2021; zero shares issued and outstanding at december 31, 2022 and 2021 | ||||||||||||||||||||||||||||||||||||
preferred stock, 0.001 par value per share; 10,000,000 shares authorized at september 30, 2022 and december 31, 2021; zero shares issued and outstanding at september 30, 2022 and december 31, 2021 | ||||||||||||||||||||||||||||||||||||
preferred stock, 0.001 par value per share; 10,000,000 shares authorized at june 30, 2022 and december 31, 2021; zero shares issued and outstanding at june 30, 2022 and december 31, 2021 | ||||||||||||||||||||||||||||||||||||
preferred stock, 0.001 par value per share; 10,000,000 shares authorized at march 31, 2022 and december 31, 2021; zero shares issued and outstanding at march 31, 2022 and december 31, 2021 | ||||||||||||||||||||||||||||||||||||
preferred stock, 0.001 par value per share; 10,000,000 shares authorized at december 31, 2021 and 2020; zero shares issued and outstanding at december 31, 2021 and 2020 | ||||||||||||||||||||||||||||||||||||
convertible preferred stock, 0.001 par value per share; 10,000,000 shares authorized at september 30, 2021 and december 31, 2020; zero shares issued and outstanding at september 30, 2021 and december 31, 2020 | ||||||||||||||||||||||||||||||||||||
convertible preferred stock, 0.001 par value per share; 10,000,000 shares authorized at june 30, 2021 and december 31, 2020; zero shares issued and outstanding at june 30, 2021 and december 31, 2020 | ||||||||||||||||||||||||||||||||||||
restricted cash, current | 132,000 | 132,000 | 132,000 | |||||||||||||||||||||||||||||||||
preferred stock, 0.001 par value per share; 10,000,000 shares authorized at december 31, 2020 and 2019; zero shares issued and outstanding at december 31, 2020 and 2019 | ||||||||||||||||||||||||||||||||||||
deferred revenue | 344,000 | 1,435,000 | 2,069,000 | 2,108,000 | 240,000 | 277,000 | 405,000 | |||||||||||||||||||||||||||||
convertible preferred stock, 0.001 par value per share; 10,000,000 shares authorized at december 31, 2019 and 2018; zero shares issued and outstanding at december 31, 2019 and 2018 | ||||||||||||||||||||||||||||||||||||
restricted cash | 386,000 | 386,000 | 132,000 | 132,000 | 132,000 | 132,000 | 132,000 | 132,000 | ||||||||||||||||||||||||||||
deferred rent, current portion | 86,000 | 79,000 | 73,000 | 66,000 | 48,000 | 44,000 | ||||||||||||||||||||||||||||||
operating lease liability | 670,000 | |||||||||||||||||||||||||||||||||||
intangible assets | 1,453,000 | 1,453,000 | 1,453,000 | |||||||||||||||||||||||||||||||||
liabilities, convertible preferred stock, and stockholders’ equity | ||||||||||||||||||||||||||||||||||||
deferred rent, long-term portion | 26,000 | 47,000 | 67,000 | 66,000 | 81,000 | |||||||||||||||||||||||||||||||
deferred tax liability | 253,000 | 305,000 | 509,000 | |||||||||||||||||||||||||||||||||
convertible preferred stock, 0.001 par value per share; 10,000,000 shares authorized at december 31, 2018 and 2017; zero shares issued and outstanding at december 31, 2018 and 2017 | ||||||||||||||||||||||||||||||||||||
total liabilities, convertible preferred stock, and stockholders’ equity | 54,873,000 | 64,508,000 | 73,124,000 | 80,699,000 | 90,786,000 | |||||||||||||||||||||||||||||||
convertible preferred stock, 0.001 par value per share; 10,000,000 shares authorized at september 30, 2018 and december 31, 2017; zero shares issued and outstanding at september 30, 2018 and december 31, 2017 | ||||||||||||||||||||||||||||||||||||
convertible preferred stock, 0.001 par value per share; 10,000,000 shares authorized at june 30, 2018 and december 31, 2017; zero shares issued and outstanding at june 30, 2018 and december 31, 2017 | ||||||||||||||||||||||||||||||||||||
convertible preferred stock, 0.001 par value per share; 10,000,000 shares authorized at march 31, 2018 and december 31, 2017; zero shares issued and outstanding at march 31, 2018 and december 31, 2017 | ||||||||||||||||||||||||||||||||||||
liabilities, convertible preferred stock and stockholders' equity | ||||||||||||||||||||||||||||||||||||
income taxes payable | 67,000 | |||||||||||||||||||||||||||||||||||
convertible preferred stock, 0.001 par value, 10,000,000 shares authorized and zero shares issued and outstanding at december 31, 2017; 0.0001 par value, 22,081,852 shares authorized and 4,311,770 shares issued and outstanding at december 31, 2016; aggregate liquidation preference of zero and 11,583 at december 31, 2017 and 2016, respectively | ||||||||||||||||||||||||||||||||||||
stockholders' equity | ||||||||||||||||||||||||||||||||||||
total stockholders' equity | 78,917,000 | 55,436,000 | ||||||||||||||||||||||||||||||||||
total liabilities, redeemable convertible preferred stock and stockholders' equity | 85,222,000 | |||||||||||||||||||||||||||||||||||
liabilities, convertible preferred stock, and stockholders’ deficit | ||||||||||||||||||||||||||||||||||||
convertible preferred stock | ||||||||||||||||||||||||||||||||||||
stockholders’ deficit: | ||||||||||||||||||||||||||||||||||||
marketable securities | 21,949,000 | 39,745,000 | 50,019,000 | 58,318,000 | 64,182,000 | 62,263,000 | 35,096,000 | |||||||||||||||||||||||||||||
property and equipment and other assets | 18,000 | 72,000 | 312,000 | 351,000 | 224,000 | 223,000 | 195,000 | 157,000 | ||||||||||||||||||||||||||||
accrued direct program expenses | 8,000 | 87,000 | 2,837,000 | 2,742,000 | 2,609,000 | 1,555,000 | 1,457,000 | 1,682,000 | ||||||||||||||||||||||||||||
accrued restructuring charges | 2,584,000 | 2,251,000 | ||||||||||||||||||||||||||||||||||
accrued employee benefits | 146,000 | 119,000 | 1,345,000 | 985,000 | 712,000 | 1,675,000 | 1,336,000 | 977,000 | ||||||||||||||||||||||||||||
accrued other liabilities | 25,000 | 54,000 | 44,000 | 36,000 | 61,000 | 195,000 | 120,000 | 32,000 | ||||||||||||||||||||||||||||
preferred stock, 0.001 par value; 10,000,000 shares authorized and no shares issued and outstanding for both periods presented | ||||||||||||||||||||||||||||||||||||
cash, cash equivalents and marketable securities | 61,035,000 | |||||||||||||||||||||||||||||||||||
working capital | 55,164,000 | |||||||||||||||||||||||||||||||||||
convertible preferred stock with liquidation preference; 0.001 par value; zero and 23,228,986 shares authorized, respectively; zero and 19,978,986 shares issued and outstanding, respectively | ||||||||||||||||||||||||||||||||||||
stockholders’ equity | ||||||||||||||||||||||||||||||||||||
preferred stock, 0.001 par value; 10,000,000 and zero shares authorized, respectively; no shares issued and outstanding |
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